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If you look at the PD survey, you'll see that there are actually many programs in these fields that *consider* DOs.
For these programs, it's something like 60-75% never rank DOs, 25-30% rarely rank DOs, and 0-10% often rank DOs.
This means that approximately 1/3 of such programs will actually consider a DO's application.
But consider doesn't mean rank highly. For that, you need to actually be competitive with the MDs. Obviously board scores, but it seems like the real hurdle for a DO is a) the research and b) the LORs from people who's words actually hold weight.
I mean, I think a DO with a) a 250+ step 1, b) 2-4 solid pubs in the field, and c) LORs from people that matter, will probably have a decent shot of matching one of these fields in ACGME. But the number of DOs with all of these are extremely small... and the number of DOs willing to skip the AOA match even smaller.
The other issue here is that these fields all have very low #s of PGY-1 or PGY-2 spots (for non-advanced ones). An average program in any one of the aforementioned fields has 3-5 residents/yr, and most heavily recruit from home students and rotators so if you end up lower than #10 or 15 on a PDs ranklist, there's no way in hell you're matching there. Even if you pass all the hurdles you mentioned, you still have to usurp X number of MDs on the ranklist to have a decent shot at matching.